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First, as a co-factor for key enzymes and a signaling molecule, Cu 2+ directly promotes rapid tumor cell proliferation and metabolic reprogramming by activating pathways such as MAPK and PI3K/Akt while stabilizing HIF-1α, establishing a state of “copper addiction.” Second, copper drives the HIF-1α/VEGF axis and activates lysyl oxidase (LOX), thereby promoting neovascularization and extracellular matrix remodeling, which supplies nutrients to the tumor and paves the way for metastasis