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Once lipid peroxides are formed, they propagate the peroxidation to the neighboring PUFAs-containing phospholipids with the presence of ferrous iron, and produce more lipid hydroxyl and peroxyl radicals, unless they are rapidly neutralized ( Figure 1 ) ( Alternatively, enzymatic lipid peroxidation is mediated by various iron-dependent enzymes ( 2.3 Antioxidant defense systems Antioxidant defense systems play a critical role in counteracting ferroptotic stress in cell