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This was observed after a frataxin minigene (made up of FXN exon1, CMV promoter, part of intron 1 and the entire exon 2) transfected into COS7 cells and HeLa cells showed splicing deficiency when the GAA dominant strand was transcribed (Baralle et al., 2008)

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[44] Similarly, genetic variations in iron transporter genes known to increase serum iron levels also reduce lifespan and the average number of years spent in good health